Principles of Evolution

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When Darwin first developed his hypothesis of descent with modification, he used a short list of basic evolutionary mechanisms or principles to explain how it worked.

These principles are helpful in understanding how organisms can change over time to become distinct species.

Below I have outlined Darwin’s original principles, as well as a number of other principles I have developed to help guide my own thinking about evolutionary topics.

If you come from a religious background, or have simply never spent much time considering the theory of evolution, understanding these principles can help you overcome the mental constructs or gaps that may be causing difficulty in understanding the process of evolution.

Darwin’s Original Principles

Below are the evolutionary principles that Charles Darwin originally outlined in his scientifically groundbreaking book The Origin of Species.

1. Competition

Within each generation, species produce more individuals than can survive within their given environment. Individuals must then compete within their own species for natural resources, survival, and the chance to pass on their genes. This competition reveals the traits most well adapted for survival.

2. Phenotypic Variation / Heritable Differences

Heritable, genetic differences can be found within the individuals of each species. These differences manifest themselves as both visible and invisible traits and create advantages or disadvantages for the individual. From the perspective of a species, variation is preferred and necessary, as it provides a higher chance of survival for that species.

3. Survival of the Best Adapted

The individuals with the genetic characteristics best suited for survival within the environment will survive and pass on their heritable genes. Advantages manifest themselves in any number of characteristics or traits — strength, speed, endurance, intelligence, social skills, resistance to pathogens, etc…

4. Descent with Modification / Phylogenesis

Reproductive isolation of groups, within a species, leads to the divergence of genetic characteristics, which allows new species to emerge from a common ancestor. The line of descent from one common ancestor, to the various species that have arisen from it, is known as the Phylogenetic tree.

Variation is a feature of natural populations and every population produces more progeny than its environment can manage. The consequences of this overproduction is that those individuals with the best genetic fitness for the environment will produce offspring that can more successfully compete in that environment. Thus the subsequent generation will have a higher representation of these offspring and the population will have evolved.

Charles Darwin

Additional Principles

In addition to the principles laid out by Darwin, there are a number of other principles that I have come across, which help me understand how evolution has and is taking place.

Below I use my own words to explain these principles, yet I do not take credit for them. I have learned all I know about evolution from the work of others.

5. Absence of Design / Intelligent Influence

This is a big one, so let’s start here. Most religious dogmas provide the idea that some intelligence or living being designed, created, and guides the universe in a state of consistent equilibrium. This belief provides a wealth of emotional support for those who ponder difficult, existential questions such as how life is possible and what the purpose of existence might be.

However, to fully understand evolution it is necessary to let go of the idea of design, purpose, and intelligent influence. When you are able to let go of these constructs many things come into focus — such as the flaws in our own biology and psychology, the genetic connections we have with other species, the power religion and nationalism have over us, and the mass extinctions to life that have taken place over time.

And, the most direct reason for dismissing the idea of intelligent design is the lack of substantive evidence for the existence of an intelligent designer.

“The universe is so unhuman, that is, it goes its way with so little thought of man. He is but an incident, not an end. We must adjust our notions to the discovery that things are not shaped to him, but that he is shaped to them. The air was not made for his lungs, but he has lungs because there is air; the light was not created for his eye, but he has eyes because there is light. All the forces of nature are going their own way; man avails himself of them, or catches a ride as best he can. If he keeps his seat, he prospers; if he misses his hold and falls, he is crushed.

John Burroughs

Nothing in biology makes sense except in the light of evolution.

Theodosius Dobzhansky

Evolution is not based on design. It has no agenda and no end goal. Species simply move forward in relation to their changing environments.

Understanding the lack of design in the universe also opens us up to understand the additional principles I have laid out below.

6. Evolution Cannot Produce Perfection

As humans we tend to see ourselves as the pinnacle of existence, the perfect species for which the Earth and the Universe were created. We may sit at the top from an intellectual and even food chain perspective, yet this in no way indicates that we are the center of the universe. Nor that we have achieved perfection as a species. Nor that we are the pinnacle of evolution.

There is no perfection, there is no pinnacle. The best evolution can produce is forward movement. This means that evolution can never go back to fix mistakes. Any weakness in genetic programming that we inherited from our ancestors moves forward with and beyond us.

This is in stark contrast to the technology we produce as humans. In intelligent design, such as can be seen in our phones, cars, and medicines. Mistakes or weaknesses can be removed and improved upon. Evolution is not making conscious decisions, so it cannot go back and start over.

Weaknesses in design, as long as they are not detrimental to survival, will always be with us.

We are one of many appearances of the thing called Life; we are not its perfect image, for it has no perfect image except Life, and life is multitudinous and emergent in the stream of time.

Loren Eiseley

Natural selection is a blind and undirected consequence of the interaction between variation and the environment. Natural selection exists only in the continuous present of the natural world: it has no memory of its previous actions, no plans for the future, or underlying purpose.

Henry Gee

Despite the Great Chain of Being’s traditional ranking of humans between animals and angels, there is no evolutionary justification for the common assumption that evolution is somehow ‘aimed’ at humans, or that humans are ‘evolution’s last word.

Richard Dawkins

7. Survival of the Good Enough

Building upon the two previous principles, this principle helps us to see how the process of Natural Selection takes place just a bit clearer.

I came across the phrase “survival of the good enough” in a book by the popular television character Bill Nye, in his book Undeniable: Evolution and the Science of Creation.

What this principle means is that again, perfection is not required for survival and is thus not the end goal of evolution. Species do not require the advancement of every single characteristic to survive. We have yet to discover a species that is evolved to survive each and every situation.

In order to survive, individuals within a species must only possess enough advantages characteristics to survive long enough within a given environment to pass on their genes.

When we refer to “survival of the fittest”, we should understand that fittest really means “best adapted” or “most well adapted”, and not “perfectly adapted”.

The biggest boys on the block at any given time aren’t necessarily the fittest, or the most efficient, and the game isn’t over. The game is never over; there’s no finish line this side of heat death. And so, neither can there be any winners. There are only those who haven’t yet lost.

Peter Watts

8. Evolutionary Change Occurs at the Genetic Level

Our genes can be thought of as a sequence of codes that determine the traits we possess. Thus, every visible characteristic of plants, and animals is based on the DNA sequence found in the nucleus of the cells.

This is the level at which evolutionary change occurs. If an individual is to have the advantage of being bigger, faster, stronger, or smarter it will be the genes of that individual that determines this advantage.

When a change takes place within a species, it is due to changes in the gene sequence of the individual. Thus, we do not see radical changes within the evolutionary process.

Oftentimes mutations take place within the DNA of an individual. Whether these mutations are beneficial or detrimental they will be passed on to the individuals offspring.

Over time, seemingly small mutations can change the traits of an entire species, or even lead to the descent of a new species within a genus.

Evolution is not a genetically controlled distortion of one adult form into another; it is a genetically controlled alteration in a developmental program.

Richard Dawkins

9. Simple on Top of Simple Becomes Complex

The small changes that take place at the genetic level are simple, but over time these simple changes can lead to more complex organic structures.

These complex results take time (see the next principle), yet when we take the time to dwell upon them, the possible path of change reveals itself.

We see this same process take place in the world of technology, only at a much more rapid pace. Think about a car and all of the small iterations that have taken place over time to allow us to produce the amazing vehicles we have today.

These vehicles are the result of small discoveries and tweaks over time. Evolution is a similar process of trial and error, only it happens independent of a guide or desired outcome. And again, evolution can not go back and scrap an old design for a new one.

If it could be demonstrated that any complex organ existed, which could not possibly have been formed by numerous, successive, slight modifications, my theory would absolutely break down. But I can find no such case.

Charles Darwin

The world doesn’t change in front of your eyes, it changes behind your back.

Terry Hayes

10. Evolution Takes Time (a lot of time)

To arrive at the world we see today, with its biodiversity and ecological diversity, the process of evolution needed a lot of time — nearly 3.6 billion years.

Our Homo sapiens brains are programmed to focus on matters of immediacy. What will we eat today? Where will we sleep tonight? What dangers do we need to watch out for in our immediate environment?

Thus, the deep time that has been required for evolution is incomprehensible to the human brain. The average lifespan of a modern American is roughly 79 years. This amount of time from an evolutionary perspective is insignificant.

A billion years is equal to 1,000 million years, and even a million years is incomprehensible. The geological evidence suggests that earth is roughly 4.5 billion years old, or 4,500 million years, or 58,227,848 human life spans. Just try to imagine that family tree.

To put this into perspective, consider an analogy sited by professor of Psychology, William Von Hipple, in his book the Social Leap. If the entire existence of the Earth was a single year (one Earth Year), then life would not have appeared until June, creatures that possessed heads wold not have appeared until October, and humans would not have appeared until nearly the end of December 31st.

Thus, life takes time.

Parting Thoughts

These principles have helped me to conceptualize the process of evolution, as well as answer questions I’ve had related to my own psychology, the social interactions I experience, and the biological world I see around me.

More-a-more, I see how these principles can be used to conceptualize where we’ve arrived in regards to our humorously human ideologies, beliefs, institutions, cultures,  and politics.
They are a key to understanding the present, through information about the past. And just maybe, they can be a gateway toward a successful future.

A loving heart is the truest wisdom.

Charles Dickens


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